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Plant Science Today (PST)

Review Articles

Vol. 13 No. 3 (2026)

Sustainably advancing plant physiology through chitosan nanoformulations

DOI
https://doi.org/10.14719/pst.13552
Submitted
6 January 2026
Published
10-07-2026 — Updated on 25-09-2026
Versions

Abstract

Modern agriculture faces significant challenges due to excessive agrochemical use, resulting in environmental degradation and reduced sustainability. Bionanomaterials have emerged as eco-friendly alternatives, among which chitosan is widely recognised for its biodegradability, biocompatibility and multifunctional properties. This review demonstrates findings from recent studies (2015–25), highlighting that conventional nutrient use efficiency remains below 50 % for macronutrients and < 5 % for micronutrients, whereas chitosan-based nanomaterials (NMs) significantly enhance nutrient delivery and utilisation. These NMs function as nanofertilisers, nanocarriers, biostimulants and nanopesticides, improving nutrient uptake, enzymatic activity, stress tolerance and overall plant growth. From a physiological perspective, they enable controlled release, targeted delivery and modulation of metabolic processes, thereby enhancing crop productivity while reducing agrochemical dependence. However, key challenges persist, including limited field-scale validation, lack of long-term environmental safety data and absence of standardised formulations. Future research should focus on large-scale validation, mechanistic insights and integration with precision agriculture. Overall, chitosan-based nanotechnology offers a promising and sustainable strategy for advancing plant physiology, though its successful field application requires further validation and standardisation.

References

  1. 1. National Academies of Sciences, Engineering, and Medicine. Science Breakthroughs 2030: A Strategy for Food and Agricultural Research. Washington (DC): The National Academies Press; 2019.
  2. 2. Pathak VM, Verma VK, Rawat BS, Kaur B, Babu N, Sharma A, et al. Current status of pesticide effects on environment, human health and its eco-friendly management as bioremediation: A comprehensive review. Front Microbiol. 2022;13:962619. https://doi.org/10.3389/fmicb.2022.962619
  3. 3. Ahmad Z, Tahseen S, Wasi A, Ganie IB, Shahzad A, Emamverdian A, et al. Nanotechnological interventions in agriculture. Nanomaterials. 2022;12:2667. https://doi.org/10.3390/nano12152667
  4. 4. Gabal E, Chatterjee S, Ahmed FK, Abd-Elsalam KA. Carbon nanomaterial applications in air pollution remediation. In: Carbon Nanomaterials for Agri-Food and Environmental Applications. Amsterdam: Elsevier; 2020. p. 133–53. https://doi.org/10.1016/B978-0-12-819786-8.00007-4
  5. 5. Saleem H, Zaidi SJ, Ismail AF, Goh PS. Advances of nanomaterials for air pollution remediation and their impacts on the environment. Chemosphere. 2022;287:132083. https://doi.org/10.1016/j.chemosphere.2021.132083
  6. 6. Pramanik B, Sar P, Bharti R, Gupta RK, Purkayastha S, Sinha S, et al. Multifactorial role of nanoparticles in alleviating environmental stresses for sustainable crop production and protection. Plant Physiol Biochem. 2023;201:107831. https://doi.org/10.1016/j.plaphy.2023.107831
  7. 7. Karamchandani BM, Dalvi SG, Bagayatkar M, Banat IM, Satpute SK. Prospective of chitosan and chitosan-based nanoparticles formulations in sustainable agricultural practices. Biocatal Agric Biotechnol. 2024;58:103210. https://doi.org/10.1016/j.bcab.2024.103210
  8. 8. Chookhongkha N, Rattanasena P, Chamswarng C. Effect of chitosan on seed quality of chilli (Capsicum annuum L.) during storage. Acta Hortic. 2013;973:231–36. https://doi.org/10.17660/ActaHortic.2013.973.32
  9. 9. Szemruch CL, Vila Cova CG, García FA, Astiz Gassó MM, Sanchez SE, Colinas A. Optimizing biopolymer seed coatings: Impact of sodium alginate and chitosan on the viability of Trichoderma harzianum, Bradyrhizobium japonicum and soybean seed quality. Int J Agric Environ Sci. 2024;11(6):1–8. https://doi.org/10.14445/23942568/IJAES-V11I6P101
  10. 10. Skrzypczak C, Górski A, Tylkowski B. Canola seed coating with formulations based on sodium alginate, chitosan, and Trichoderma spp. J Plant Prot Res. 2021;61(3):308–16.
  11. 11. García AI, González R, Reyes C, Silva M. Effect of chitosan film coating on the germination and early growth of sesame (Sesamum indicum L.). Agronomy. 2022;12(3):666. https://doi.org/10.3390/agronomy12030666
  12. 12. Çetin Karaca A, Cinfir A. Effects of biopolymer-based seed coating on bacterial viability and plant growth in Phaseolus vulgaris L. Int J Agric Nat Res. 2023;6(2):108–15.
  13. 13. Vijaykumar M, Sudhakar S, Sai Ram R, Geetha N. Development of dual microbe-based double-layered biopolymer seed coating with Trichoderma harzianum and Bradyrhizobium sp. for groundnut. Int J Biol Macromol. 2024;252:129876.
  14. 14. Chandrika KS, Sudhakar S, Sai Ram R, Geetha N. Chitosan-based biopolymer seed coating with Trichoderma harzianum improves germination, disease suppression, and yield in oilseed crops. Appl Microbiol Biotechnol. 2024.
  15. 15. Ahmed EM. Hydrogel: Preparation, characterization, and applications: A review. Carbohydr Polym. 2014;112:1–10.
  16. 16. Prakash V, Chauhan D, Meena RS. Controlled release of nitrogen through nano-chitosan encapsulation improves soil fertility and plant growth. Ecotoxicol Environ Saf. 2018;149:498–506.
  17. 17. Singh R, Yadav R, Verma A. Use of chitosan as a biopolymer for improving soil fertility and crop productivity. Soil Use Manag. 2019;35(2):245–52.
  18. 18. Farouk S, Albarakaty FM, Al-Mutairi KA, Elsayed FA, El-Ghany TMA. Potential of chitosan nanoparticles for improving growth and productivity of tomato under greenhouse conditions. Sci Rep. 2020;10:20299.
  19. 19. El-Saadony MT, Desoky ESM, Saad AM, Eid RS, Selem E, Rady MM. Chitosan nanoparticles alleviate salt stress in plants: A review on mechanisms and applications. Front Plant Sci. 2021;12:696972.
  20. 20. Khalifa RKHM, El-Kafafi SH, Hassan HA, Fattah MA. Effect of chitosan biopolymer matrix enriched with micronutrients on tomato growth and soil health. J Plant Nutr. 2022;45(12):1850–62.
  21. 21. Khan MM, Shakeel M, Rehman A, Saeed MA, Abid M, Aslam MM, et al. Chitosan bead-infused biochar improves the soil physico-chemical properties and productivity of sunflower irrigated with wastewater. Acta Physiol Plant. 2024;46:69.
  22. 22. Iqbal DN, Tariq Z, Philips B, Sadiqa A, Ahmad M, Al-Ahmary KM, et al. Nanocellulose/wood ash-reinforced starch-chitosan hydrogel composites for soil conditioning and their impact on pea plant growth. RSC Adv. 2024;14:8652–64. https://doi.org/10.1039/D3RA08725E
  23. 23. Gornik K, Grzesik M, Duda BR. The effect of chitosan on rooting of grapevine cuttings and on subsequent plant growth under drought and temperature stress. J Fruit Ornam Plant Res. 2008;16:333–43.
  24. 24. Mondal M, Md Puteh A, Ismail M, Naher L. Effect of foliar application of chitosan on growth and yield in okra. Aust J Crop Sci. 2012;6(5):918–21.
  25. 25. Divya K, Thampi M, Vijayan S, Shabanamol S, Jisha MS. Chitosan nanoparticles as a rice growth promoter: Evaluation of biological activity. Arch Microbiol. 2022;204(1):95. https://doi.org/10.1007/s00203-021-02669-w
  26. 26. Mawale KS, Nandini B, Giridhar P. Copper and silver nanoparticle seed priming and foliar spray modulate plant growth and thrips infestation in Capsicum spp. ACS Omega. 2024;9(3):3430–44. https://doi.org/10.1021/acsomega.3c06961
  27. 27. Sreethu MJ, Venugopal V. Chitosan nanoparticles as biocompatible tool in agriculture. Int J Environ Clim Chang. 2025;15(2):123–37. https://doi.org/10.9734/ijecc/2025/v15i24717
  28. 28. Ramírez-Rodríguez SC, Preciado-Rangel P, Fuente MC-DL, González-Morales S, Ortega-Ortiz H. Chitosan nanoparticles as biostimulant in lettuce (Lactuca sativa L.) plants. Phyton. 2024;93(4):777–87. https://doi.org/10.32604/phyton.2024.048096
  29. 29. Apostolova EL, Misra AN. Alterations in structural organization affect the functional ability of photosynthetic apparatus. In: Pessarakli M, editor. Handbook of Plant and Crop Physiology. 3rd ed. Boca Raton (FL): CRC Press; 2014. p. 103–20. https://doi.org/10.1201/b16675-5
  30. 30. Araus JL, Sanchez-Bragado R, Vicente R. Improving crop yield and resilience through optimization of photosynthesis: Panacea or pipe dream? J Exp Bot. 2021;72(11):3936–55. https://doi.org/10.1093/jxb/erab097
  31. 31. Mishra AN. Chlorophyll fluorescence: A practical approach to study ecophysiology of green plants. In: Adela M, Sánchez-Moreiras AM, Reigosa MJ, editors. Switzerland: Springer International Publishing; 2018. p. 77–79. https://doi.org/10.1007/978-3-319-93233-0_5
  32. 32. Stefanov M, Biswal AK, Misra M, Misra AN, Apostolova EL. Responses of photosynthetic apparatus to salt stress: Structure, function, and protection. Boca Raton (FL): CRC Press; 2019. p. 233–50. https://doi.org/10.1201/9781351104609-13
  33. 33. Moustafa-Farag M, Mohamed HI, Mahmoud A, Elkelish A, Misra AN, Guy KM, et al. Salicylic acid stimulates antioxidant defense and osmolyte metabolism to alleviate oxidative stress in watermelons under excess boron. Plants. 2020;9(6):724. https://doi.org/10.3390/plants9060724
  34. 34. Sharma G, Kumar A, Devi KA, Prajapati D, Bhagat D, Pal A, et al. Chitosan nanofertilizer to foster source activity in maize. Int J Biol Macromol. 2020;145:226–34. https://doi.org/10.1016/j.ijbiomac.2019.12.155
  35. 35. Prajapati D, Pal A, Dimkpa C, Harish, Singh U, Devi KA, et al. Chitosan nanomaterials: A prelim of next-generation fertilizers; existing and future prospects. Carbohydr Polym. 2022;288:119356. https://doi.org/10.1016/j.carbpol.2022.119356
  36. 36. Sharma G, Saharan V, Pal A, Sharma SS. Chitosan nanofertilizer to strengthen sink strength and provide resistance against PFSR (Post-Flowering Stalk Rot) disease in maize. Biocatal Agric Biotechnol. 2024;60:103303. https://doi.org/10.1016/j.bcab.2024.103303
  37. 37. Kumaraswamy RV, Saharan V, Kumari S, Choudhary RC, Pal A, Sharma SS, et al. Chitosan-silicon nanofertilizer to enhance plant growth and yield in maize (Zea mays L.). Plant Physiol Biochem. 2021;159:53–66. https://doi.org/10.1016/j.plaphy.2020.11.054
  38. 38. Kadam PM, Prajapati D, Kumaraswamy RV, Kumari S, Devi KA, Pal A, et al. Physio-biochemical responses of wheat plant towards salicylic acid-chitosan nanoparticles. Plant Physiol Biochem. 2021;162:699–705. https://doi.org/10.1016/j.plaphy.2021.03.021
  39. 39. Nandini T, Sudhalakshmi C, Sivakumar K, Parameswari E, Thangamani C. A review-chitosan nanoparticles towards enhancing nutrient use efficiency in crops. Int J Biol Macromol. 2025;141433. https://doi.org/10.1016/j.ijbiomac.2025.141433
  40. 40. Choudhary RC, Kumaraswamy RV, Kumari S, Sharma SS, Pal A, Raliya R, et al. Cu-chitosan nanoparticle boost defense responses and plant growth in maize (Zea mays L.). Sci Rep. 2017;7:9754. https://doi.org/10.1038/s41598-017-08571-0
  41. 41. Baligar VC, Fageria NK. Nutrient use efficiency in plants: an overview. In: Rakshit A, Singh HB, Sen A, editors. Nutrient Use Efficiency: From Basics to Advances. Dordrecht: Springer; 2015. p. 1–14. https://doi.org/10.1007/978-81-322-2169-2_1
  42. 42. Monreal CM, DeRosa M, Mallubhotla SC, Bindraban PS, Dimkpa C. Nanotechnologies for increasing the crop use efficiency of fertilizer-micronutrients. Biol Fertil Soils. 2016;52(3):423–37. https://doi.org/10.1007/s00374-015-1073-5
  43. 43. Duhan JS, Kumar R, Kumar N, Kaur P, Nehra K, Duhan S. Nanotechnology: the new perspective in precision agriculture. Biotechnol Rep. 2017;125:11–23. https://doi.org/10.1016/j.btre.2017.03.002
  44. 44. Gogos A, Knauer K, Bucheli TD. Nanomaterials in plant protection and fertilization: current state, foreseen applications, and research priorities. J Agric Food Chem. 2012;60(39):9781–92. https://doi.org/10.1021/jf302154y
  45. 45. Karthik MU, Maheswari. Smart fertilizer strategy for better crop production. Agric Rev. 2021;42(1):12–21. https://doi.org/10.18805/ag.R-1877
  46. 46. Brunetti C, Fini A. Fertilization in urban landscape. In: Routledge Handbook of Urban Forestry. Oxfordshire: Routledge; 2017. p. 434–48. https://doi.org/10.4324/9781315627106-29
  47. 47. Liu G, Zotarelli L, Li Y, Dinkins D, Wang Q, Ozores-Hampton M. Controlled-release and slow-release fertilizers as nutrient management tools. Gainesville (FL): University of Florida, IFAS Extension Service, US Department of Agriculture; 2014. https://doi.org/10.32473/edis-hs1255-2014
  48. 48. Timilsena YP, Adhikari R, Casey P, Muster T, Gill H, Adhikari B. Enhanced efficiency fertilisers: a review of formulation and nutrient release patterns. J Sci Food Agric. 2015;95(6):1131–42. https://doi.org/10.1002/jsfa.6812
  49. 49. Shifren A. A local solution for a global problem: technology-forcing municipal ordinances to promote enhanced efficiency fertilizers. Colum J Envt L. 2022;47:146. https://doi.org/10.52214/cjel.v47i1.9130
  50. 50. Li T, Gao B, Tong Z, Yang Y, Li Y. Chitosan and graphene oxide nanocomposites as coatings for controlled-release fertilizer. Water Air Soil Pollut. 2019;230:1–9. https://doi.org/10.1007/s11270-019-4173-2
  51. 51. Kumaraswamy RV, Kumari S, Choudhary RC, Sharma SS, Pal A, Raliya R, et al. Salicylic acid functionalized chitosan nanoparticle: A sustainable biostimulant for plant. Int J Biol Macromol. 2019;123:59–69. https://doi.org/10.1016/j.ijbiomac.2018.10.202
  52. 52. Gohari G, Jiang M, Manganaris GA, Zhou J, Fotopoulos V. Next generation chemical priming: with a little help from our nanocarrier friends. Trends Plant Sci. 2024;29(2):150–66. https://doi.org/10.1016/j.tplants.2023.11.024
  53. 53. Nair R. Effects of nanoparticles on plant growth and development. In: Kole C, Kumar D, Khodakovskaya M, editors. Plant Nanotechnology. Cham: Springer; 2016. p. 95–118. https://doi.org/10.1007/978-3-319-42154-4_5
  54. 54. Siddiqi KS, Husen A. Fabrication of metal nanoparticles from fungi and metal salts: scope and application. Nanoscale Res Lett. 2016;11:98. https://doi.org/10.1186/s11671-016-1311-2
  55. 55. Antoniou J, Liu F, Majeed H, Qi J, Yokoyama W, Zhong F. Physicochemical and morphological properties of size-controlled chitosan-tripolyphosphate nanoparticles. Colloids Surf A Physicochem Eng Asp. 2015;465:137–46. https://doi.org/10.1016/j.colsurfa.2014.10.040
  56. 56. Choudhary RC, Kumaraswamy RV, Kumari S, Sharma SS, Pal A, Raliya R, et al. Zinc encapsulated chitosan nanoparticle to promote maize crop yield. Int J Biol Macromol. 2019;127:126–35. https://doi.org/10.1016/j.ijbiomac.2018.12.274
  57. 57. Aazami MA, Maryam M, Farzad, Gohari G. Protective effects of chitosan based salicylic acid nanocomposite (CS-SA NCs) in grape (Vitis vinifera cv. ‘Sultana’) under salinity stress. Sci Rep. 2023;13:883. https://doi.org/10.1038/s41598-023-27618-z
  58. 58. Gohari G, Farhadi H, Panahirad S, Zareei E, Labib P, Jafari H, et al. Mitigation of salinity impact in spearmint plants through the application of engineered chitosan-melatonin nanoparticles. Int J Biol Macromol. 2023;224:893–907. https://doi.org/10.1016/j.ijbiomac.2022.10.175
  59. 59. Panahirad S, Dadpour M, Gohari G, Akbari A, Mahdavinia G, Jafari H, et al. Putrescine-functionalized carbon quantum dot (put-CQD) nanoparticle: A promising stress protecting agent against cadmium stress in grapevine (Vitis vinifera cv. Sultana). Plant Physiol Biochem. 2023;197:107653. https://doi.org/10.1016/j.plaphy.2023.107653
  60. 60. Sheikhalipour M, Mohammadi SA, Esmaielpour B, Spanos A, Mahmoudi R, Mahdavinia GR, et al. Seedling nanopriming with selenium-chitosan nanoparticles mitigates the adverse effects of salt stress by inducing multiple defence pathways in bitter melon plants. Int J Biol Macromol. 2023;242:124923. https://doi.org/10.1016/j.ijbiomac.2023.124923
  61. 61. dos Santos Guaraldo MM, Pereira TM, dos Santos HO, de Oliveira TL, Pereira WVS, Von Pinho EVDR. Priming with sodium nitroprusside and hydrogen peroxide increases cotton seed tolerance to salinity and water deficit during seed germination and seedling development. Environ Exp Bot. 2023;209:105294. https://doi.org/10.1016/j.envexpbot.2023.105294
  62. 62. Giglou MT, Giglou RH, Esmaeilpour B, Azarmi R, Padash A, Falakian M, et al. A new method in mitigation of drought stress by chitosan-coated iron oxide nanoparticles and growth stimulant in peppermint. Ind Crops Prod. 2022;187:115286. https://doi.org/10.1016/j.indcrop.2022.115286
  63. 63. Corradini E, de Moura MR, Mattoso LHC. A preliminary study of the incorporation of NPK fertilizer into chitosan nanoparticles. Express Polym Lett. 2010;4(8):509–15. https://doi.org/10.3144/expresspolymlett.2010.64
  64. 64. Pereira AES, Narciso AM, Seabra AB, Fraceto LF. Evaluation of the effects of nitric oxide-releasing nanoparticles on plants. J Phys Conf Ser. 2014;617:012025. Proceedings of the 4th International Conference on Safe Production and Use of Nanomaterials (Nanosafe2014); 2014; Grenoble, France. https://doi.org/10.1088/1742-6596/617/1/012025
  65. 65. Abdel-Aziz HMM, Hasaneen MNA, Omer AM. Nano chitosan-NPK fertilizer enhances the growth and productivity of wheat plants grown in sandy soil. Span J Agric Res. 2016;14(1):e0902. https://doi.org/10.5424/sjar/2016141-8205
  66. 66. Ha NMC, Nguyen TH, Wang S-L, Nguyen AD. Preparation of NPK nanofertilizer based on chitosan nanoparticles and its effect on biophysical characteristics and growth of coffee in greenhouse. Res Chem Intermed. 2019;45:51–63. https://doi.org/10.1007/s11164-018-3630-7
  67. 67. Marchiol L, Iafisco M, Fellet G, Adamiano A. Nanotechnology support the next agricultural revolution: Perspectives to enhancement of nutrient use efficiency. Adv Agron. 2020;161:27–116. https://doi.org/10.1016/bs.agron.2019.12.001
  68. 68. Smith EN, van Aalst M, Tosens T, Niinemets Ü, Stich B, Morosinotto T, et al. Improving photosynthetic efficiency toward food security: Strategies, advances, and perspectives. Mol Plant. 2023;16(10):1547–63. https://doi.org/10.1016/j.molp.2023.08.017
  69. 69. Yin H, Bai XF, Du YG. The primary study of oligochitosan inducing resistance to Sclerotinia sclerotiorum on B. napus. J Biotechnol. 2008;136:600–1. https://doi.org/10.1016/j.jbiotec.2008.07.1217
  70. 70. Guan YJ, Hu J, Wang XJ, Shao CX. Seed priming with chitosan improves maize germination and seedling growth in relation to physiological changes under low temperature stress. J Zhejiang Univ Sci B. 2009;10(6):427–33. https://doi.org/10.1631/jzus.B0820373
  71. 71. El-Miniawy SM, Ragab ME, Youssef SM, Metwally AA. Response of strawberry plants to foliar spraying of chitosan. Res J Agric Biol Sci. 2013;9(6):366–72.
  72. 72. Chamnanmanoontham N, Pongprayoon W, Pichayangkura R, Roytrakul S. S. Chitosan enhances rice seedling growth via gene expression network between nucleus and chloroplast. Plant Growth Regul. 2015;75(1):101–14. https://doi.org/10.1007/s10725-014-9935-7
  73. 73. Behboudi F, Tahmasebi Z, Sarvestani MZ, Kassaee SAM, Modares S, Sorooshzadeh A, et al. Evaluation of chitosan nanoparticles effects on yield and yield components of barley (Hordeum vulgare L.) under late season drought stress. J Water Environ Nanotechnol. 2018;3(1):22–39.
  74. 74. Iriti M, Sironi M, Gomarasca S, Casazza AP, Soave C, Faoro F. Cell death mediated antiviral activity of chitosan. Plant Physiol Biochem. 2006;44:893–900. https://doi.org/10.1016/j.plaphy.2006.10.009
  75. 75. Iriti M, Faoro F. Chitosan as a MAMP searching for PRR. Plant Signal Behav. 2009;4:66–8. https://doi.org/10.4161/psb.4.1.7408
  76. 76. Kumaraswamy RV, Kumari S, Choudhary RC, Pal A, Raliya R, Biswas P, et al. Engineered chitosan based nanomaterials: Bioactivities, mechanisms and perspectives in plant protection and growth. Int J Biol Macromol. 2018;113:494–506. https://doi.org/10.1016/j.ijbiomac.2018.02.130
  77. 77. Hadwiger LA. Multiple effects of chitosan on plant systems: Solid science or hype. Plant Sci. 2013;208:42–9. https://doi.org/10.1016/j.plantsci.2013.03.007
  78. 78. Kumar A, Prajapati D, Devi KA, Pal A, Choudhary U, Dashora A, et al. Slow-release Zn application through Zn-chitosan nanoparticles in wheat to intensify source activity and sink strength. Plant Physiol Biochem. 2021;168:272–81. https://doi.org/10.1016/j.plaphy.2021.10.013
  79. 79. Saharan V, Kumaraswamy RV, Choudhary RC, Kumari S, Pal A, Raliya R, et al. Cu-chitosan nanoparticle mediated sustainable approach to enhance seedling growth in maize by mobilizing reserved food. J Agric Food Chem. 2016;64(31):6148–55. https://doi.org/10.1021/acs.jafc.6b02239
  80. 80. Sahu S, Gautam S, Singh A, Lohani P, Sharma C, Pathak P, et al. Synthesis and characterization of chitosan-zinc-salicylic acid nanoparticles: A plant biostimulant. Int J Biol Macromol. 2023;253(8):127602. https://doi.org/10.1016/j.ijbiomac.2023.127602
  81. 81. Chen A, Peng C, Su C, Ma Y, Zhan X, Chen J, et al. Chitosan-copper hybrid nanoflowers: A novel nanopesticide for controlling Rhizoctonia solani infection in crops. J Agric Food Chem. 2024;72(41):22545–53. https://doi.org/10.1021/acs.jafc.4c06345
  82. 82. Masoumi Z, Haghighi M, Mozafarian M. Effects of foliar spraying with melatonin and chitosan nano-encapsulated melatonin on tomato (Lycopersicon esculentum L. cv. Falcato) plants under salinity stress. BMC Plant Biol. 2024;24:961. https://doi.org/10.1186/s12870-024-05672-7
  83. 83. Seyed-Hajizadeh H, Faraji-Chelanolya A, Zahedi SM, Moghadam A, Mahdavinia G, Kaya O. Nanochitosan-encapsulated melatonin: An eco-friendly strategy to delay petal senescence in cut gerbera flowers. BMC Plant Biol. 2024;24(1):1024. https://doi.org/10.1186/s12870-024-05725-x
  84. 84. Fregonezi B, Pereira AES, Ferreira JM, Fraceto LF, Gomes DG, Oliveira HC. Seed priming with nanoencapsulated gibberellic acid triggers beneficial morphophysiological and biochemical responses of tomato plants under different water conditions. Agronomy. 2024;14(3):588. https://doi.org/10.3390/agronomy14030588
  85. 85. Azimi F, Oraei M, Gohari G, Panahirad S, Farmarzi A. Chitosan-selenium nanoparticles (Cs-Se NPs) modulate the photosynthesis parameters, antioxidant enzymes activities and essential oils in Dracocephalum moldavica L. under cadmium toxicity stress. Plant Physiol Biochem. 2021;167:257–68. https://doi.org/10.1016/j.plaphy.2021.08.013
  86. 86. González ME, Pérez LA, Rodríguez FJ. A novel biostimulant from chitosan nanoparticles and microalgae extract enhances tomato growth and yield. Sci Hortic. 2023;310:111234.
  87. 87. Methela NJ, Pande A, Islam MS, Yun BW. Chitosan-GSNO nanoparticles: A positive modulator of drought stress tolerance in soybean. BMC Plant Biol. 2023;23:639. https://doi.org/10.1186/s12870-023-04640-x
  88. 88. Kumar R, Sharma P, Singh S. Role of chitosan nanoparticles in enhancing growth and yield of toria (Brassica campestris). J Agric Sci Technol. 2025;27(3):123–34.
  89. 89. Zheng Q, Wu J, Yan W, Zhu S, Miao X, Wang R, et al. Green synthesis of a chlorfenapyr chitosan nanopesticide for maize root application: Reducing environmental pollution and risks to nontarget organisms. Int J Biol Macromol. 2023;253(4):126988. https://doi.org/10.1016/j.ijbiomac.2023.126988
  90. 90. Wang X, Tarahomi M, Sheibani R, Xia C, Wang W. Progresses in lignin, cellulose, starch, chitosan, chitin, alginate, and gum/carbon nanotube (nano)composites for environmental applications: A review. Int J Biol Macromol. 2023;241:124472. https://doi.org/10.1016/j.ijbiomac.2023.124472
  91. 91. Egorov AR, Kirichuk AA, Rubanik VV, Rubanik VV, Tskhovrebov AG, Kritchenkov AS. Chitosan and its derivatives: Preparation and antibacterial properties. Materials (Basel). 2023;16(18):6076. https://doi.org/10.3390/ma16186076
  92. 92. Yang Y, Aghbashlo M, Gupta VK, Amiri H, Pan J, Tabatabaei M, et al. Chitosan nanocarriers containing essential oils as a green strategy to improve the functional properties of chitosan: A review. Int J Biol Macromol. 2023;236:123954. https://doi.org/10.1016/j.ijbiomac.2023.123954
  93. 93. Fan Z, Wang L, Qin Y, Li P. Activity of chitin/chitosan/chitosan oligosaccharide against plant pathogenic nematodes and potential modes of application in agriculture: A review. Carbohydr Polym. 2023;306:120592. https://doi.org/10.1016/j.carbpol.2023.120592
  94. 94. Karamchandani BM, Dalvi SG, Bagayatkar M, Banat IM, Satpute SK. Prospective applications of chitosan and chitosan-based nanoparticles formulations in sustainable agricultural practices. Biocatal Agric Biotechnol. 2024;58:103210. https://doi.org/10.1016/j.bcab.2024.103210
  95. 95. Román-Doval R, Torres-Arellanes SP, Tenorio-Barajas AY, Gómez-Sánchez A, Valencia-Lazcano AA. Chitosan: Properties and its application in agriculture in context of molecular weight. Polymers. 2023;15(13):2867. https://doi.org/10.3390/polym15132867
  96. 96. Wu J, Chang J, Liu J, Huang J, Song Z, Xie X, et al. Chitosan-based nanopesticides enhanced anti-fungal activity against strawberry anthracnose as 'sugar-coated bombs'. Int J Biol Macromol. 2023;253(4):126947. https://doi.org/10.1016/j.ijbiomac.2023.126947
  97. 97. Darwesh OM, Abdelmaksoud NM, El-Bakry AM. Nano-chitosan as a bio-enhancer for improving insecticide formulations: A study on indoxacarb and methomyl against cotton leafworm. Int J Biol Macromol. 2024;282(6):137465. https://doi.org/10.1016/j.ijbiomac.2024.137465
  98. 98. Abdel-Nasser A, Fathy HM, Badr AN, Barakat OS, Hathout AS. Chitosan nanoparticles loaded with Lactobacillus rhamnosus bioactive metabolites: Preparation, characterization, and antifungal activity. Heliyon. 2025;11(2):e41875. https://doi.org/10.1016/j.heliyon.2025.e41875
  99. 99. Abenaim L, Conti B. Chitosan as a control tool for insect pest management: A review. Insects. 2023;14(12):949. https://doi.org/10.3390/insects14120949

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