Skip to main navigation menu Skip to main content Skip to site footer

Review Articles

Vol. 13 No. sp1 (2026): Recent Advances in Agriculture

Recent advances in the alleviation of salt stress in plants

DOI
https://doi.org/10.14719/pst.10723
Submitted
18 July 2025
Published
24-03-2026

Abstract

Salt stress is one of the crucial abiotic factors limiting plant growth and production. It is caused by high salt levels in the soil, which limit the availability of irrigated water and other minerals required for plant growth. It can cause poor crop yields and food insecurity, particularly in coastal and desert areas. Mitigating plant salt stress is critical for increasing agricultural yields and maintaining food security. There are numerous approaches to reducing the negative consequences of salt stress, including genetic modification, hormone therapy  and improved soil management practices. Recent discoveries in the field include microbial consortia, nanotechnology, metabolomics, systems biology  and CRISPR-Cas technologies for improving plant salt tolerance. Minimising salt stress in plants is crucial for enhancing agricultural output and food security and several quantitative methods can be utilised to accomplish this goal. This review will address recent findings on salt tolerance in plants, utilising microbial, morpho-physical, biochemical  and genetic approaches  and nanoparticle applications.

References

  1. 1. Bruce TJA. Tackling the threat to food security caused by crop pests in the new millennium. Food Secur. 2010;2(2):133–41. https://doi.org/10.1007/s12571-010-0061-8
  2. 2. Newton AC, Johnson SN, Gregory PJ. Implications of climate change for diseases, crop yields and food security. Euphytica. 2011;179:3–18 https://doi.org/10.1007/s10681-011-0359-4
  3. 3. Enenkel M, See L, Bonifacio R, Boken V, Chaney N, Vinck P, et al. Drought and food security-improving decision-support via new technologies and innovative collaboration. Glob Food Secur. 2015;4:51–5. https://doi.org/10.1016/j.gfs.2014.08.005
  4. 4. Kumar P, Sharma PK. Soil salinity and food security in India. Front Sustain Food Syst. 2020;4:533781. https://doi.org/10.3389/fsufs.2020.533781
  5. 5. Reed C, Anderson W, Kruczkiewicz A, Nakamura J, Gallo D, Seager R, et al. The impact of flooding on food security across Africa. Proc Natl Acad Sci USA. 2022;119 (43):e2119399119.
  6. 6. Prasad SS. Enzyme dynamics in aerobic plants under flood stress (review). Biologiya. 2025;60(5). https://doi.org/10.15389/agrobiology.2025.5.792eng
  7. 7. Prasad SS, Kurubar AR, Hugar A, Ramesh G, Umesh MR, Meena MK. Cost effectiveness of gladiolus production under drip fertigation and planting geometry. Indian J Hortic. 2022;79(3):353–62. https://doi.org/10.5958/0974-0112.2022.00048.2
  8. 8. Singh G, Bundela DS, Sethi M, Lal K, Kamra SK. Remote sensing and geographic information system for appraisal of salt-affected soils in India. J Environ Qual. 2010;39:5–15. https://doi.org/10.2134/jeq2009.0032
  9. 9. Hossain A, Pamanick B, Venugopalan VK, Ibrahimova U, Rahman MA, Siyal AL, et al. Emerging roles of plant growth regulators for plants adaptation to abiotic stress–induced oxidative stress. In: Aftab T, Naeem M, editors. Emerging plant growth regulators in agriculture. Cambridge (MA): Academic Press; 2022. p. 1–72. https://doi.org/10.1016/B978-0-323-91005-7.00010-2
  10. 10. Medina N, Rouhi Rad M, Nozari S, Suter JF, Bailey RT, Sahoo D. Water scarcity compounds the negative effects of salinity on irrigated agriculture. Water Resour Res. 2025;61. https://doi.org/10.1029/2024WR038504
  11. 11. Cetin M, Kirda C. Spatial and temporal changes of soil salinity in a cotton field irrigated with low-quality water. J Hydrol. 2003;273(1–4):238–49. https://doi.org/10.1016/S0022-1694 (02)00268-8
  12. 12. Rhoades JD, Krueger DB, Reed MJ. The effect of soil-mineral weathering on the sodium hazard of irrigation waters. Soil Sci Soc Am J. 1968;32:643–77. https://doi.org/10.2136/sssaj1968.03615995003200050020x
  13. 13. Nassar IN, Horton R. Salinity and compaction effects on soil water evaporation and water and solute distributions. Soil Sci Soc Am J. 1999;63:752–8. https://doi.org/10.2136/sssaj1999.634752x
  14. 14. Sun J, Yang G, Zhang W, Zhang Y. Effects of heterogeneous salinity on growth, water uptake and tissue ion concentrations of alfalfa. Plant Soil. 2016;408:211–26. https://doi.org/10.1007/s11104-016-2922-1
  15. 15. Pearcy RW, Ustin SL. Effects of salinity on growth and photosynthesis of three California tidal marsh species. Oecologia. 1984;62:68–73. https://doi.org/10.1007/BF00377375
  16. 16. Grattan SR, Grieve CM. Salinity-mineral nutrient relations in horticultural crops. Sci Hortic. 1998;78:127–57. https://doi.org/10.1016/S0304-4238 (98)00192-7
  17. 17. Mukhtar S, Mirza BS, Mehnaz S, Mirza MS, Mclean J, Malik KA. Impact of soil salinity on the microbial structure of halophyte rhizosphere microbiome. World J Microbiol Biotechnol. 2018;34 (9):136. https://doi.org/10.1007/s11274-018-2509-5
  18. 18. Sheldon AR, Dalal RC, Kirchhof G, Kopittke PM, Menzies NW. The effect of salinity on plant-available water. Plant Soil. 2017;418:477–91. https://doi.org/10.1007/s11104-017-3309-7
  19. 19. Tripathi S, Tiwari K, Mahra S, Victoria J, Rana S, Tripathi DK, et al. Nanoparticles and root traits: mineral nutrition, stress tolerance and interaction with rhizosphere microbiota. Planta. 2024;259:112. https://doi.org/10.1007/s00425-024-04409-y
  20. 20. Sha S, Wang G, Liu J, Wang M, Wang L, Liu Y, et al. Regulation of photosynthetic function and reactive oxygen species metabolism in sugar beet (Beta vulgaris L.) cultivars under waterlogging stress and associated tolerance mechanisms. Plant Physiol Biochem. 2024;211:108651. https://doi.org/10.1016/j.plaphy.2024.108651
  21. 21. Gangaiah B, Yadav AK. Modern crop management practices for pearl millet cultivation in Asia. In: Tonapi VA, Thirunavukkarasu N, Gupta S, Gangashetty PI, Yadav O, editors. Pearl Millet in the 21st Century. Singapore: Springer; 2024. p. 445–72. https://doi.org/10.1007/978-981-99-5890-0_18
  22. 22. Yang Y, Guo Y. Unraveling salt stress signaling in plants. J Integr Plant Biol. 2018;60:796–804. https://doi.org/10.1111/jipb.12689
  23. 23. Wang CF, Han GL, Yang ZR, Li YX, Wang BS. Plant salinity sensors: current understanding and future directions. Front Plant Sci. 2022;13:859224. https://doi.org/10.3389/fpls.2022.859224
  24. 24. Boussora F, Triki T, Bennani L, Bagues M, Ben Ali S, Ferchichi A, et al. Mineral accumulation, relative water content and gas exchange are the main physiological regulating mechanisms to cope with salt stress in barley. Sci Rep. 2024;14:14840. https://doi.org/10.1038/s41598-024-65967-5
  25. 25. Rasheed Y, Khalid F, Ashraf H, Asif K, Maqsood MF, Naz N, et al. Enhancing plant stress resilience with osmolytes and nanoparticles. J Soil Sci Plant Nutr. 2024;24:1871–1906. https://doi.org/10.1007/s42729-024-01821-x
  26. 26. Habibi N, Tayobong RRP, Parneel, Terada N, Sanada A, Koshio K. Novel insights into seed priming for tomato plants: restoring root vitality in the face of salt stress. Hortic Environ Biotechnol. 2025;66:361–80. https://doi.org/10.1007/s13580-024-00651-1
  27. 27. Munns R, Tester M. Mechanisms of salinity tolerance. Annu Rev Plant Biol. 2008;59:651–81. https://doi.org/10.1146/annurev.arplant.59.032607.092911
  28. 28. Zahra N, Al Hinai MS, Hafeez MB, Rehman A, Wahid A, Siddique KHM, et al. Regulation of photosynthesis under salt stress and associated tolerance mechanisms. Plant Physiol Biochem. 2022;178:55–69. https://doi.org/10.1016/j.plaphy.2022.03.003
  29. 29. Parida AK, Das AB. Salt tolerance and salinity effects on plants: a review. Ecotoxicol Environ Saf. 2005;60(3):324–49. https://doi.org/10.1016/j.ecoenv.2004.06.010
  30. 30. Seshagiri S, Varma A, Tallapragada P. Piriformospora indica-mediated amelioration of salt stress in Solanum melongena L. plants under greenhouse-a study on stomatal opening/closure. Geomicrobiol J. 2025;42:213–23. https://doi.org/10.1080/01490451.2025.2452568
  31. 31. Başak H, Aydin A, Yetişir H, Turan M. Salt stress effects on hybrid bottle gourd (Lagenaria siceraria) rootstock candidates plant growth, hormones and nutrient content. J Crop Health. 2025;77:1–15. https://doi.org/10.1007/s10343-024-01087-4
  32. 32. Chrouqi L, Sammama H, Azenzem R, Koussa T, Alfeddy MN. Impact of Bacillus thuringiensis (S48) in enhancing the oxidative stress defence systems of two wheat varieties against salt stress. Russ J Plant Physiol. 2025;72:1–12. https://doi.org/10.1134/S1021443725600059
  33. 33. Ajila-Celi GE, Lata-Tenesaca LF, Calzada KP, de Cassia Alves R, da Cruz MCP, Junior JSP, et al. Exogenous ascorbic acid mitigates salt-induced damage in soybean by modulating photosynthesis, antioxidant defense and ionic homeostasis. Acta Physiol Plant. 2025;47:1–14. https://doi.org/10.1007/s11738-025-03770-z
  34. 34. Kumari R, Rakhra G, Alsahli AA, Bhat JA, Ahmad P. Exploring the potential of signalling molecules hydrogen sulfide and nitric oxide in augmenting salt stress resilience in bitter gourd. BMC Plant Biol. 2025;25:42. https://doi.org/10.1186/s12870-025-06942-8
  35. 35. Kumari R, Khan MN, Parrey ZA, Kapoor P, Mir BA, Taziun T, et al. Synergistic effects of hydrogen sulfide and nitric oxide in enhancing salt stress tolerance in cucumber seedlings. Physiol Plant. 2025;177:e7010. https://doi.org/10.1111/ppl.70109
  36. 36. Goel D, Singh AK, Yadav V, Babbar SB, Bansal KC. Overexpression of osmotin gene confers tolerance to salt and drought stresses in transgenic tomato (Solanum lycopersicum L.). Protoplasma. 2010;245:133–41. https://doi.org/10.1007/s00709-010-0158-0
  37. 37. Sherpa MT, Sharma L, Bag N, Das S. Isolation, characterization and evaluation of native rhizobacterial consortia developed from the rhizosphere of rice grown in organic state Sikkim, India and their effect on plant growth. Front Microbiol. 2021;12:713660. https://doi.org/10.3389/fmicb.2021.713660
  38. 38. Quesada V, García-Martínez S, Piqueras P, Ponce MR, Micol JL. Genetic architecture of NaCl tolerance in Arabidopsis. Plant Physiol. 2002;130:951–63. https://doi.org/10.1104/pp.006536
  39. 39. Dermendjiev G, Schnurer M, Weiszmann J, Wilfinger S, Ott E, Gebert C, et al. Tissue-specific proteome and subcellular microscopic analyses reveal the effect of high salt concentration on actin cytoskeleton and vacuolization in aleurone cells during early germination of barley. Int J Mol Sci. 2021;22(17):9642. https://doi.org/10.3390/ijms22179642
  40. 40. Witzel K, Weidner A, Surabhi GK, Varshney RK, Kunze G, Buck-Sorlin GH, et al. Comparative analysis of the grain proteome fraction in barley genotypes with contrasting salinity tolerance during germination. Plant Cell Environ. 2010;33:211–22. https://doi.org/10.1111/j.1365-3040.2009.02071.x
  41. 41. Akbari M, Karbor S, Afshar A, Ferguson L, Farajpour M, Dillard T, et al. The role of rootstock selection in pistachio cultivation. Horticulturae. 2024;10(12):1278. https://doi.org/10.3390/horticulturae10121278
  42. 42. Li X, Xu Y, Zhang J, Xu K, Zheng X, Luo J, et al. Integrative physiology and transcriptome reveal salt-tolerance differences between two licorice species: ion transport, Casparian strip formation and flavonoids biosynthesis. BMC Plant Biol. 2024;24:391. https://doi.org/10.1186/s12870-024-04911-1
  43. 43. Xie X, Gan L, Wang C, He T. Salt-tolerant plant growth-promoting bacteria as a versatile tool for combating salt stress in crop plants. Arch Microbiol. 2024;206:312. https://doi.org/10.1007/s00203-024-04071-8
  44. 44. Nadeem SM, Zahir ZA, Naveed M, Asghar HN, Arshad M. Rhizobacteria capable of producing ACC-deaminase may mitigate salt stress in wheat. Soil Sci Soc Am J. 2010;74:533–42. https://doi.org/10.2136/sssaj2008.0240
  45. 45. Sofy MR, Aboseidah AA, Heneidak SA, Ahmed HR. ACC deaminase-containing endophytic bacteria ameliorate salt stress in Pisum sativum through reduced oxidative damage and induction of antioxidative defense systems. Environ Sci Pollut Res. 2021;28:40971–91. https://doi.org/10.1007/s11356-021-13585-3
  46. 46. Dias MC, Silva S, Galhano C, Lorenzo P. Olive tree belowground microbiota: plant growth-promoting bacteria and fungi. Plants. 2024;13(13):1848. https://doi.org/10.3390/plants13131848
  47. 47. Yan J, Smith MD, Glick BR, Liang Y. Effects of ACC deaminase containing rhizobacteria on plant growth and expression of Toc GTPases in tomato (Solanum lycopersicum) under salt stress. Botany. 2014;92:775–81. https://doi.org/10.1139/cjb-2014-0038
  48. 48. Sukweenadhi J, Balusamy SR, Kim YJ, Lee CH, Kim YJ, Koh SC, et al. A growth-promoting bacteria, Paenibacillus yonginensis DCY84T enhanced salt stress tolerance by activating defense-related systems in Panax ginseng. Front Plant Sci. 2018;9:813. https://doi.org/10.3389/fpls.2018.00813
  49. 49. Sharma K, Sharma S, Vaishnav A, Jain R, Singh D, Singh HB, et al. Salt-tolerant PGPR strain Priestia endophytica SK1 promotes fenugreek growth under salt stress by inducing nitrogen assimilation and secondary metabolites. J Appl Microbiol. 2022;133:2802–13. https://doi.org/10.1111/jam.15735
  50. 50. Acharya BR, Gill SP, Kaundal A, Sandhu D. Strategies for combating plant salinity stress: the potential of plant growth-promoting microorganisms. Front Plant Sci. 2024;15:1406913. https://doi.org/10.3389/fpls.2024.1406913
  51. 51. Siddika A, Rashid AA, Khan SN, Khatun A, Karim MM, Prasad PVV, et al. Harnessing plant growth-promoting rhizobacteria, Bacillus subtilis and Bacillus aryabhattai to combat salt stress in rice: a study on the regulation of antioxidant defense, ion homeostasis and photosynthetic parameters. Front Plant Sci. 2024;15:1419764. https://doi.org/10.3389/fpls.2024.1419764
  52. 52. Pan J, Peng F, Xue X, You Q, Zhang W, Wang T, et al. The growth promotion of two salt-tolerant plant groups with PGPR inoculation: a meta-analysis. Sustainability. 2019;11:20378. https://doi.org/10.3390/su11020378
  53. 53. Galicia-Campos E, García-Villaraco Velasco A, Montero-Palmero MB, Gutiérrez-Mañero FJ, Ramos-Solano B. Modulation of photosynthesis and ROS scavenging response by beneficial bacteria in Olea europaea plantlets under salt stress conditions. Plants. 2022;11(20):2748. https://doi.org/10.3390/plants11202748
  54. 54. Ji J, Zhang J, Wang X, Song W, Ma B, Wang R, et al. The alleviation of salt stress on rice through increasing photosynthetic capacity, maintaining redox homeostasis and regulating soil enzyme activities by Enterobacter sp. JIV1 assisted with putrescine. Microbiol Res. 2024;278:127590. https://doi.org/10.1016/j.micres.2023.127590
  55. 55. Gohari G, Jiang M, Manganaris GA, Zhou J, Fotopoulos V. Next generation chemical priming: with a little help from our nanocarrier friends. Trends Plant Sci. 2024;29(4):464–77. https://doi.org/10.1016/j.tplants.2023.11.024
  56. 56. Riaz N, Yousaf Z, Younas A, Hyder S, Aftab A, Maqbool Z, et al. Microbiomes: a road map of medicinal plant growth, disease control and abiotic stress management. In: Symbiotic association of microorganisms with medicinal and herbal plants. Boca Raton: CRC Press; 2024. p. 97–116. https://doi.org/10.1201/b23155-6
  57. 57. Anand G, Goel V, Dubey S, Sharma S. Tailoring the rhizospheric microbiome of Vigna radiata by adaptation to salt stress. Plant Growth Regul. 2021;93(1):79–88. https://doi.org/10.1007/s10725-020-00667-4
  58. 58. Patel M, Islam S, Husain FM, Yadav VK, Park HK, Yadav KK, et al. Bacillus subtilis ER-08, a multifunctional plant growth-promoting rhizobacterium, promotes the growth of fenugreek (Trigonella foenum-graecum L.) plants under salt and drought stress. Front Microbiol. 2023;14:1208743. https://doi.org/10.3389/fmicb.2023.1208743
  59. 59. Orozco-Mosqueda MC, Glick BR, Santoyo G. ACC deaminase in plant growth-promoting bacteria (PGPB): an efficient mechanism to counter salt stress in crops. Microbiol Res. 2020;235:126439. https://doi.org/10.1016/j.micres.2020.126439
  60. 60. Mayak S, Tirosh T, Glick BR. Plant growth-promoting bacteria confer resistance in tomato plants to salt stress. Plant Physiol Biochem. 2004;42:565–72. https://doi.org/10.1016/j.plaphy.2004.05.009
  61. 61. El-Ramady H, Prokisch J, Mansour H, Bayoumi YA, Shalaby TA, Veres S, et al. Review of crop response to soil salinity stress: possible approaches from leaching to nano-management. Soil Syst. 2024;8 (1):11. https://doi.org/10.3390/soilsystems8010011
  62. 62. El Yamani M, Cordovilla MP. Tolerance mechanisms of olive tree (Olea europaea) under saline conditions. Plants. 2024;13:2094. https://doi.org/10.3390/plants13152094
  63. 63. Wagan S, Ali M, Khoso MA, Alam I, Dinislam K, Hussain A, et al. Deciphering the role of WRKY transcription factors in plant resilience to alkaline salt stress. Plant Stress. 2024;13:100526. https://doi.org/10.1016/j.stress.2024.100526
  64. 64. Hadizadeh I, Peivastegan B, Nielsen KL, Auvinen P, Sipari N, Pirhonen M. Transcriptome analysis unravels the biocontrol mechanism of Serratia plymuthica A30 against potato soft rot caused by Dickeya solani. PLoS One. 2024;19(11):e0308744. https://doi.org/10.1371/journal.pone.0308744
  65. 65. Guan Q, Tan B, Kelley TM, Tian J, Chen S. Physiological changes in Mesembryanthemum crystallinum during the C3 to CAM transition induced by salt stress. Front Plant Sci. 2020;11:283. https://doi.org/10.3389/fpls.2020.00283
  66. 66. Ahmad B, Mukarram M, Choudhary S, Petrík P, Dar TA, Khan MMA. Adaptive responses of nitric oxide (NO) and its intricate dialogue with phytohormones during salinity stress. Plant Physiol Biochem. 2024;211:108504. https://doi.org/10.1016/j.plaphy.2024.108504
  67. 67. Atul S, Pradeep KS, Shiva SP, Satendra S, Pragati M, Rajeshwar JR, et al. Differential effects of high-temperature stress on the morpho-physiological traits of different wheat (Triticum aestivum L.) genotypes. Plant Sci Today. 2025;12(1):1–10. https://doi.org/10.14719/pst.6103
  68. 68. Adishesha K, Umesh BC, Jeevitha D, Sujatha HT, A, Prasad SS, et al. Temperature regime impacts on reproductive performance in various chickpea (Cicer arietinum L.) genotypes. Indian J Agric Res. 2025;59:1–7. https://doi.org/10.18805/IJARe.A-6367
  69. 69. Tariq F, Ma C, Zhao S. Integrative dynamics of cell wall architecture and plant growth under salt stress. Front Plant Sci. 2025;16:1644412. https://doi.org/10.3389/fpls.2025.1644412
  70. 70. Iqbal U, Daad A, Ali A, Gul MF, Aslam MU, Rehman FU, et al. Surviving the desert’s grasp: decipherment of phreatophyte Tamarix aphylla (L.) Karst. adaptive strategies for arid resilience. Plant Sci. 2024;348:112201. https://doi.org/10.1016/j.plantsci.2024.112201
  71. 71. Yagi N, Fujita S, Nakamura M. Plant microtubule nucleating apparatus and its potential signaling pathway. Curr Opin Plant Biol. 2024;81:102624. https://doi.org/10.1016/j.pbi.2024.102624
  72. 72. Sun Y, Xiao Z, Chen B, Zhao Y, Dai J. Advances in material-assisted electromagnetic neural stimulation. Adv Mater. 2024;36 (40):2400346. https://doi.org/10.1002/adma.202400346
  73. 73. Bohnert HJ, Ayoubi P, Borchert C, Bressan RA, Burnap RL, Cushman JC, et al. A genomics approach towards salt stress tolerance. Plant Physiol Biochem. 2001;39:295–311. https://doi.org/10.1016/S0981-9428 (00)01237-7
  74. 74. Zhao C, Sandhu D, Ferreira JFS. Transcript analysis of two spinach cultivars reveals the complexity of salt tolerance mechanisms. ACS Agric Sci Technol. 2021;1:64–75. https://doi.org/10.1021/acsagscitech.0c00063
  75. 75. Chen F, Fang P, Peng Y, Zeng W, Zhao X, Ding Y, et al. Comparative proteomics of salt-tolerant and salt-sensitive maize inbred lines to reveal the molecular mechanism of salt tolerance. Int J Mol Sci. 2019;20 (19):4725. https://doi.org/10.3390/ijms20194725
  76. 76. Yang D, Zhang J, Li M, Shi L. Metabolomics analysis reveals the salt-tolerant mechanism in Glycine soja. J Plant Growth Regul. 2017;36:460–71. https://doi.org/10.1007/s00344-016-9654-6
  77. 77. Ingle KP, Singh A, Sahni T, Aakanksha K, Kumar S, Geethika P, et al. Genome editing in synthetic biology for sustainable production of biomolecules. In: Biomanufacturing for sustainable production of biomolecules. Singapore: Springer Nature; 2023. p. 315–29. https://doi.org/10.1007/978-981-19-7911-8_16
  78. 78. Bogoutdinova LR, Lazareva EM, Chaban IA, Kononenko NV, Dilovarova T, Khaliluev MR, et al. Salt stress-induced structural changes are mitigated in transgenic tomato plants over-expressing superoxide dismutase. Biology. 2020;9(9):297. https://doi.org/10.3390/biology9090297
  79. 79. Yan H, Li Q, Park SC, Wang X, Liu YJ, Zhang YG, et al. Overexpression of CuZnSOD and APX enhance salt stress tolerance in sweet potato. Plant Physiol Biochem. 2016;109:20–7. https://doi.org/10.1016/j.plaphy.2016.09.003
  80. 80. Prasad SS, Singh A, Patil K, Ramteke PW, Kumar N, Shukla PK, et al. Next-generation sequencing (NGS) application in plant breeding. In: Advances in plant genomics. Cham (CH): Springer; 2024. p. 149–71. https://doi.org/10.1007/978-3-031-68586-6_6
  81. 81. Prasad SS. Enhancing the throughput of design–build–test–learn cycle for the future perspective of synthetic biology in plants (review). Sel’skokhozyaistvennaya Biologiya. 2024;59(5):831–46.
  82. 82. Zhao B, Liu Q, Luo L, Zhou H, Zhang X, Ma F, et al. Suppression of MdPRP6 enhances adaptation of apple plants to long-term drought. Physiol Plant. 2025;177:e70099. https://doi.org/10.1111/ppl.70099
  83. 83. Singh D, Debnath P, Sane AP, Sane VA. Tomato (Solanum lycopersicum) WRKY23 enhances salt and osmotic stress tolerance by modulating the ethylene and auxin pathways in transgenic Arabidopsis. Plant Physiol Biochem. 2023;195:330–40. https://doi.org/10.1016/j.plaphy.2023.01.002
  84. 84. Cao X, Ma W, Zeng F, Cheng Y, Ma Z, Mao J, et al. Grape BES1 transcription factor gene VvBES1-3 confers salt tolerance in transgenic Arabidopsis. Gene. 2023;854:147059. https://doi.org/10.1016/j.gene.2022.147059
  85. 85. Ding X, Liu B, Liu H, Sun X, Sun X, Wang W, et al. A new CIPK gene CmCIPK8 enhances salt tolerance in transgenic chrysanthemum. Sci Hortic. 2023;308:111562. https://doi.org/10.1016/j.scienta.2022.111562
  86. 86. Yu M, Liu J, Du B, Zhang M, Wang A, Zhang L. NAC transcription factor PwNAC11 activates ERD1 by interaction with ABF3 and DREB2A to enhance drought tolerance in transgenic Arabidopsis. Int J Mol Sci. 2021;22(13):6952. https://doi.org/10.3390/ijms22136952
  87. 87. Luo C, Akhtar M, Min W, Alam Y, Ma T, Shi Y, et al. The suppressed expression of a stress-responsive gene OsDSR2 enhances rice tolerance to drought and salt stress. J Plant Physiol. 2023;282:153927. https://doi.org/10.1016/j.jplph.2023.153927
  88. 88. Muthamilarasan M, Khandelwal R, Yadav CB, Bonthala VS, Khan Y, Prasad M. Identification and molecular characterization of MYB transcription factor superfamily in C4 model plant foxtail millet (Setaria italica L.). PLoS One. 2014;9(10):e109920. https://doi.org/10.1371/journal.pone.0109920
  89. 89. Lühmann KL, Seemann S, Martinek N, Ostendorp S, Kehr J. The glycine-rich domain of GRP7 plays a crucial role in binding long RNAs and facilitating phase separation. Sci Rep. 2024;14:66955. https://doi.org/10.1038/s41598-024-66955-5
  90. 90. Sun J, Manmathan H, Sun C, Peebles CAM. Examining the transcriptional response of overexpressing anthranilate synthase in hairy roots of Catharanthus roseus by RNA-seq. BMC Plant Biol. 2016;16:79. https://doi.org/10.1186/s12870-016-0794-4
  91. 91. Zhou X, Rao S, Wrightstone E, Sun T, Lui ACW, Welsch R, et al. Phytoene synthase: the key rate-limiting enzyme of carotenoid biosynthesis in plants. Front Plant Sci. 2022;13:884720. https://doi.org/10.3389/fpls.2022.884720
  92. 92. Chen Y, Li C, Zhang B, Yi J, Yang Y, Kong C, et al. The role of the late embryogenesis-abundant (LEA) protein family in development and abiotic stress response: comprehensive expression analysis of potato (Solanum tuberosum). Genes (Basel). 2019;10(2):148. https://doi.org/10.3390/genes10020148
  93. 93. Zhang SS, Sun L, Dong X, Lu SJ, Tian W, Liu JX. Cellulose synthesis genes CESA6 and CSI1 are important for salt stress tolerance in Arabidopsis. J Integr Plant Biol. 2016;58:623–6. https://doi.org/10.1111/jipb.12442
  94. 94. Qu D, Wu F, Yang J, Li M, Yang L, Xie R, et al. Transcription factor PtNAC101 negatively regulates lignin synthesis and salt tolerance in Populus trichocarpa. Environ Exp Bot. 2023;205:105149. https://doi.org/10.1016/j.envexpbot.2022.105149
  95. 95. Ren JP, Li L, Cai XL, Huang W, Chao DY, Zhu MZ, et al. A rice quantitative trait locus for salt tolerance encodes a sodium transporter. Nat Genet. 2005;37(10):1141–6. https://doi.org/10.1038/ng1643
  96. 96. Zhang X, Gong X, Yu H, Su X, Cheng S, Huang J, et al. The proline-rich protein MdPRP6 confers tolerance to salt stress in transgenic apple (Malus domestica). Sci Hortic. 2023;308:111581. https://doi.org/10.1016/j.scienta.2022.111581
  97. 97. Ren Y, Lv J, Wang H, Li L, Peng Y, Qu LJ. A comparative proteomics approach to detect unintended effects in transgenic Arabidopsis. J Genet Genomics. 2009;36:629–39. https://doi.org/10.1016/S1673-8527 (08)60155-1
  98. 98. Adarme-Vega TC, Thomas-Hall SR, Schenk PM. Towards sustainable sources for omega-3 fatty acids production. Curr Opin Biotechnol. 2014;26:14–8. https://doi.org/10.1016/j.copbio.2013.08.003
  99. 99. Yu Y, Guo DD, Min DH, Cao T, Ning L, Jiang QY, et al. Foxtail millet MYB-like transcription factor SiMYB16 confers salt tolerance in transgenic rice by regulating the phenylpropanoid pathway. Plant Physiol Biochem. 2023;195:310–21. https://doi.org/10.1016/j.plaphy.2022.11.032
  100. 100. Treutter D. Significance of flavonoids in plant resistance: a review. Environ Chem Lett. 2006;4:147–57. https://doi.org/10.1007/s10311-006-0068-8
  101. 101. Cui X, Cao Y, Zhang H, Zhang L. Picea wilsonii NAC transcription factor PwNAC1 interacts with RNA-binding protein PwRBP1 and synergistically enhances drought and salt tolerance in transgenic Arabidopsis. Environ Exp Bot. 2023;206:105174. https://doi.org/10.1016/j.envexpbot.2022.105174
  102. 102. Jiao K, Han J, Guo B, Wu Y, Zhang L, Li Y, et al. MbNAC22, a Malus baccata NAC transcription factor, increases drought and salt tolerance in Arabidopsis. Agronomy. 2023;13(5):1374. https://doi.org/10.3390/agronomy13051374
  103. 103. Huang Y, Du B, Yu M, Cao Y, Liang K, Zhang L. Picea wilsonii NAC31 and DREB2A cooperatively activate ERD1 to modulate drought resistance in transgenic Arabidopsis. Int J Mol Sci. 2024;25(4):2037. https://doi.org/10.3390/ijms25042037
  104. 104. Iqbal MZ, Liang Y, Anwar M, Fatima A, Hassan MJ, Ali A, et al. Overexpression of auxin/indole-3-acetic acid gene TrIAA27 enhances biomass, drought and salt tolerance in Arabidopsis thaliana. Plants. 2024;13(19):2684. https://doi.org/10.3390/plants13192684
  105. 105. Awere CO, Rakkammal K, Mwaura MM, Anadebe VC, Ramesh M. Hairy-root technology: a metabolic engineering tool for specialized metabolite pathway elucidation and production of secondary metabolites. Results Eng. 2024;24:102697. https://doi.org/10.1016/j.rineng.2024.102697
  106. 106. Endler A, Kesten C, Schneider R, Zhang Y, Ivakov A, Froehlich A, et al. A mechanism for sustained cellulose synthesis during salt stress. Cell. 2015;162:1353–64. https://doi.org/10.1016/j.cell.2015.08.028
  107. 107. Lou T, Lv S, Wang J, Wang D, Lin K, Zhang X, et al. Cell size and xylem differentiation regulating genes from Salicornia europaea contribute to plant salt tolerance. Plant Cell Environ. 2024;47 (11):4184–204. https://doi.org/10.1111/pce.14905
  108. 108. Chun HJ, Baek D, Cho HM, Lee SH, Jin BJ, Yun DJ, et al. Lignin biosynthesis genes play critical roles in adaptation of Arabidopsis to high-salt stress. Plant Signal Behav. 2019;14(10):e1625697. https://doi.org/10.1080/15592324.2019.1625697
  109. 109. Kim B, Cho SH, Shim Y, Yoon H, Paek NC, Kang K. Rice OsMYB9 enhances salt stress tolerance by regulating the vacuolar Na+/H+ antiporter. Crop J. 2025;13:1718–3. https://doi.org/10.1016/j.cj.2025.08.002
  110. 110. Calzone A, Cotrozzi L, Pellegrini E, Lorenzini G, Nali C, Maathuis F. Transcriptional regulation of NHX1, SOS1 and HKT1 genes in two pomegranate cultivars under moderate salt stress. Sci Hortic. 2021;288:110309. https://doi.org/10.1016/j.scienta.2021.110309
  111. 111. Ahmad P, Hashem A, Abd-Allah EF, Alqarawi AA, John R, Egamberdieva D, et al. Role of Trichoderma harzianum in mitigating NaCl stress in Indian mustard (Brassica juncea L.) through antioxidative defense system. Front Plant Sci. 2015;6:868. https://doi.org/10.3389/fpls.2015.00868
  112. 112. Van Delden SH, Nazarideljou MJ, Marcelis LFM. Nutrient solutions for Arabidopsis thaliana: a study on nutrient solution composition in hydroponic systems. Plant Methods. 2020;16:66. https://doi.org/10.1186/s13007-020-00606-4
  113. 113. Ndiaye M, Mollier A, Diouf A, Diop TA. Mycorrhizal inoculation and fertilizer microdosing interactions in pearl millet (Pennisetum glaucum) under greenhouse conditions. Front Fungal Biol. 2024;5:1448156. https://doi.org/10.3389/ffunb.2024.1448156
  114. 114. Park HS, Kazerooni EA, Kang SM, Al-Sadi AM, Lee IJ. Melatonin enhances tolerance and recovery mechanisms in Brassica juncea under saline conditions. Front Plant Sci. 2021;12:593717. https://doi.org/10.3389/fpls.2021.593717
  115. 115. Yaschenko AE, Alonso JM, Stepanova AN. Arabidopsis as a model for translational research. Plant Cell. 2025;37(5):koae065. https://doi.org/10.1093/plcell/koae065
  116. 116. Genc Y, Taylor J, Lyons G, Li Y, Cheong J, Appelbee M, et al. Bread wheat with high salinity and sodicity tolerance. Front Plant Sci. 2019;10:1280. https://doi.org/10.3389/fpls.2019.01280
  117. 117. Tian Y, Guan B, Zhou D, Yu J, Li G, Lou Y. Responses of seed germination, seedling growth and seed yield traits to seed pretreatment in maize (Zea mays L.). Sci World J. 2014;2014:834630. https://doi.org/10.1155/2014/834630
  118. 118. Zuo G, Huo J, Yang X, Mei W, Zhang R, Khan A, et al. Photosynthetic mechanisms underlying NaCl-induced salinity tolerance in rice (Oryza sativa). BMC Plant Biol. 2024;24:723. https://doi.org/10.1186/s12870-024-04723-3
  119. 119. Dabravolski SA, Isayenkov SV. The regulation of plant cell wall organisation under salt stress. Front Plant Sci. 2023;14:1118313. https://doi.org/10.3389/fpls.2023.1118313
  120. 120. Läuchli A, Grattan SR. Plant growth and development under salinity stress. In: Advances in molecular breeding toward drought and salt tolerant crops. Dordrecht: Springer; 2007. p. 1–32. https://doi.org/10.1007/978-1-4020-5578-2_1
  121. 121. Francois LE, Grieve CM, Maas EV, Lesch SM. Time of salt stress affects growth and yield components of irrigated wheat. Agron J. 1994;86:100–7. https://doi.org/10.2134/agronj1994.00021962008600010019x
  122. 122. Yang Z, Wang Y, Wei X, Zhao X, Wang B, Sui N. Transcription profiles of genes related to hormonal regulation under salt stress in sweet sorghum. Plant Mol Biol Rep. 2017;35:586–99. https://doi.org/10.1007/s11105-017-1047-x
  123. 123. Shu K, Qi Y, Chen F, Meng Y, Luo X, Shuai H, et al. Salt stress represses soybean seed germination by negatively regulating GA biosynthesis while positively mediating ABA biosynthesis. Front Plant Sci. 2017;8:1372. https://doi.org/10.3389/fpls.2017.01372
  124. 124. Choudhary P, Pramitha L, Rana S, Verma S, Aggarwal PR, Muthamilarasan M. Hormonal crosstalk in regulating salinity stress tolerance in graminaceous crops. Physiol Plant. 2021;173(4):1587–96. https://doi.org/10.1111/ppl.13558
  125. 125. Ellouzi H, Ben Slimene Debez I, Amraoui S, Rabhi M, Hanana M, Alyami NM, et al. Effect of seed priming with auxin on ROS detoxification and carbohydrate metabolism in salt-stressed maize. BMC Plant Biol. 2024;24:541. https://doi.org/10.1186/s12870-024-05413-w
  126. 126. Yuan K, Rashotte AM, Wysocka-Diller JW. ABA and GA signaling pathways interact and regulate seed germination and seedling development under salt stress. Acta Physiol Plant. 2011;33:261–71. https://doi.org/10.1007/s11738-010-0542-6
  127. 127. Medeiros BO, Silva LAS, Sarmento SN, Rosa DA, de Souza Barbosa LC, Machado M, et al. Antagonistic interactions between cytokinin and gibberellin during initial stem growth and leaf structure of royal poinciana (Delonix regia). Trees (Berl). 2024;38:1–14. https://doi.org/10.1007/s00468-024-02562-1
  128. 128. Koevoets IT, Venema JH, Elzenga JTM, Testerink C. Roots withstanding their environment: exploiting root system architecture responses to abiotic stress to improve crop tolerance. Front Plant Sci. 2016;7:1335. https://doi.org/10.3389/fpls.2016.01335
  129. 129. Ryu H, Cho YG. Plant hormones in salt stress tolerance. J Plant Biol. 2015;58:147–55. https://doi.org/10.1007/s12374-015-0103-z
  130. 130. Garrido-Vargas F, Godoy T, Tejos R, O’Brien JA. Overexpression of the auxin receptor AFB3 in Arabidopsis results in salt stress resistance and modulation of NAC4 and SZF1. Int J Mol Sci. 2020;21 (24):9528. https://doi.org/10.3390/ijms21249528
  131. 131. Roychowdhury R, Hada A, Biswas S, Mishra S, Prusty MR, Das SP, et al. Jasmonic acid (JA) in plant immune response: unravelling complex molecular mechanisms and networking of defence signalling against pathogens. J Plant Growth Regul. 2024;43:1–22. https://doi.org/10.1007/s00344-024-11264-4
  132. 132. Hazman M, Sühnel M, Schäfer S, Zumsteg J, Lesot A, Beltran F, et al. Characterization of jasmonoyl-isoleucine (JA-Ile) hormonal catabolic pathways in rice upon wounding and salt stress. Rice. 2019;12:45. https://doi.org/10.1186/s12284-019-0303-0
  133. 133. Farhangi-Abriz S, Ghassemi-Golezani K. How can salicylic acid and jasmonic acid mitigate salt toxicity in soybean plants? Ecotoxicol Environ Saf. 2018;147:1010–6. https://doi.org/10.1016/j.ecoenv.2017.09.070
  134. 134. Sadhukhan A, Prasad SS, Mitra J, Siddiqui N, Sahoo L, Kobayashi Y, et al. How do plants remember drought? Planta. 2022;256:7. https://doi.org/10.1007/s00425-022-03924-0
  135. 135. Sarkar S, Guha A, Narayanan TN, Mondal J. Zwitterionic osmolytes revive surface charges under salt stress via dual mechanisms. J Phys Chem Lett. 2022;13(24):5660–8. https://doi.org/10.1021/acs.jpclett.2c00853
  136. 136. Thabet SG, Alomari DZ, Alqudah AM. Exploring natural diversity reveals alleles to enhance antioxidant system in barley under salt stress. Plant Physiol Biochem. 2021;166:789–98. https://doi.org/10.1016/j.plaphy.2021.06.030
  137. 137. El Moukhtari A, Cabassa-Hourton C, Farissi M, Savouré A. How does proline treatment promote salt stress tolerance during crop plant development? Front Plant Sci. 2020;11:1127. https://doi.org/10.3389/fpls.2020.01127
  138. 138. Ahmad P, Latef AAA, Hashem A, Abd Allah EF, Gucel S, Tran LSP. Nitric oxide mitigates salt stress by regulating levels of osmolytes and antioxidant enzymes in chickpea. Front Plant Sci. 2016;7:347. https://doi.org/10.3389/fpls.2016.00347
  139. 139. Manai J. Role of nitric oxide in saline stress: implications on nitrogen metabolism. Afr J Plant Sci. 2012;6 (14):376–82. https://doi.org/10.5897/AJPS12.079
  140. 140. Santisree P, Adimulam SS, Sharma K, Bhatnagar-Mathur P, Sharma KK. Insights into the nitric oxide mediated stress tolerance in plants. In: Khan MIR, Khan NA, editors. Plant signaling molecules: role and regulation under stressful environments. Cambridge (MA): Academic Press; 2019. p. 385–406. https://doi.org/10.1016/B978-0-12-816451-8.00024-1
  141. 141. Javaid MH, Chen N, Yasin MU, Fan X, Neelam A, Rehman M, et al. Green-synthesized lignin nanoparticles enhance Zea mays resilience to salt stress by improving antioxidant metabolism and mitigating ultrastructural damage. Chemosphere. 2024;359:142337. https://doi.org/10.1016/j.chemosphere.2024.142337
  142. 142. Nkomo M, Badiwe M, Niekerk LA, Gokul A, Keyster M, Klein A. p-Coumaric acid differentially alters the ion-omics profile of chia shoots under salt stress. Plants. 2024;13(11):1564. https://doi.org/10.3390/plants13111564
  143. 143. Linić I, Mlinarić S, Brkljačić L, Pavlović I, Smolko A, Salopek-Sondi B. Ferulic acid and salicylic acid foliar treatments reduce short-term salt stress in Chinese cabbage by increasing phenolic compounds accumulation and photosynthetic performance. Plants. 2021;10(11):2346. https://doi.org/10.3390/plants10112346
  144. 144. Yang J, Zhang Y, Jia J, Wang C, Fu Y. Flavonoid-lignin crosstalk: engineering metabolic flux for optimised plant growth and stress resilience. Plant Cell Environ. 2025;48:8141–60. https://doi.org/10.1111/pce.70106
  145. 145. Liu S, Liu J, Wang Y, Deng F, Deng Z. Oxidative stress: signaling pathways, biological functions and disease. MedComm. 2025;6:e702. https://doi.org/10.1002/mco2.70268
  146. 146. Gao W, Xu FC, Guo DD, Zhao JR, Liu J, Guo YW, et al. Calcium-dependent protein kinases in cotton: Insights into early plant responses to salt stress. BMC Plant Biol. 2018;18:12. https://doi.org/10.1186/s12870-018-1230-8
  147. 147. Khan I, Awan SA, Rizwan M, Akram MA, Zia-ur-Rehman M, Wang X, et al. Physiological and transcriptome analyses demonstrate the silver nanoparticles mediated alleviation of salt stress in pearl millet (Pennisetum glaucum L.). Environ Pollut. 2023;318:120863. https://doi.org/10.1016/j.envpol.2022.120863
  148. 148. Zahedi SM, Abolhassani M, Hadian-Deljou M, Feyzi H, Akbari A, Rasouli F, et al. Proline-functionalized graphene oxide nanoparticles (GO-Pro NPs): A new engineered nanoparticle to ameliorate salinity stress on grape (Vitis vinifera L. cv Sultana). Plant Stress. 2023;7:100128. https://doi.org/10.1016/j.stress.2022.100128
  149. 149. Gohari G, Panahirad S, Mohammadi A, Kulak M, Dadpour MR, Lighvan ZM, et al. Characterization of octa-aminopropyl polyhedral oligomeric silsesquioxanes (OA-POSS) nanoparticles and their effect on sweet basil (Ocimum basilicum L.) response to salinity stress. Plant Physiol Biochem. 2023;196:89–102. https://doi.org/10.1016/j.plaphy.2023.01.019
  150. 150. Song Y, Zheng C, Basnet R, Li S, Chen J, Jiang M. Astaxanthin synthesized gold nanoparticles enhance salt stress tolerance in rice by enhancing tetrapyrrole biosynthesis and scavenging reactive oxygen species in vitro. Plant Stress. 2022;6:100122. https://doi.org/10.1016/j.stress.2022.100122
  151. 151. Rossi L, Zhang W, Lombardini L, Ma X. The impact of cerium oxide nanoparticles on the salt stress responses of Brassica napus L. Environ Pollut. 2016;219:28–36. https://doi.org/10.1016/j.envpol.2016.09.060
  152. 152. Mukarram M, Petrik P, Mushtaq Z, Khan MMA, Gulfishan M, Lux A. Silicon nanoparticles in higher plants: Uptake, action, stress tolerance and crosstalk with phytohormones, antioxidants and other signalling molecules. Environ Pollut. 2022;310:119855. https://doi.org/10.1016/j.envpol.2022.119855
  153. 153. Ahsan M, Radicetti E, Jamal A, Ali HM, Sajid M, Manan A, et al. Silicon nanoparticles and indole butyric acid positively regulate the growth performance of Freesia refracta by ameliorating oxidative stress under chromium toxicity. Front Plant Sci. 2024;15:1437276. https://doi.org/10.3389/fpls.2024.1437276
  154. 154. Rizwan A, Zia-ur-Rehman M, Rizwan M, Usman M, Anayatullah S, Areej, et al. Effects of silicon nanoparticles and conventional Si amendments on growth and nutrient accumulation by maize (Zea mays L.) grown in saline-sodic soil. Environ Res. 2023;227:115740. https://doi.org/10.1016/j.envres.2023.115740
  155. 155. Siddiqi KS, Husen A, Zahra N, Moheman A. Harnessing silicon nanoparticles and various forms of silicon for enhanced plant growth performance under salinity stress: Application and mechanism. Discov Nano. 2025;20:42. https://doi.org/10.1186/s11671-025-04270-2
  156. 156. Shahpari M, Mohammadi Torkashvand A, Ghanbari Jahromi M. Silicon nanoparticles-modulated salinity stress in sage (Salvia officinalis L.) plants through optimizing biochemical status and essential oil yield. J Plant Nutr. 2026;49:522–35. https://doi.org/10.1080/01904167.2025.2558891
  157. 157. Ibrahim EA. Seed priming to alleviate salinity stress in germinating seeds. J Plant Physiol. 2016;192:38–46. https://doi.org/10.1016/j.jplph.2015.12.011
  158. 158. Avestan S, Ghasemnezhad M, Esfahani M, Byrt CS. Application of nano-silicon dioxide improves salt stress tolerance in strawberry plants. Agronomy. 2019;9(5):246. https://doi.org/10.3390/agronomy9050246
  159. 159. Khan M, Jannat A, Munir F, Fatima N, Amir R. Biochemical and molecular mechanisms of abiotic stress tolerance. In: Hasanuzzaman M, editor. Plant ecophysiology and adaptation under climate change: Mechanisms and perspectives II. Singapore: Springer; 2020. p. 187–230. https://doi.org/10.1007/978-981-15-2172-0_9
  160. 160. Peña-Calzada K, Olivera-Viciedo D, Calero-Hurtado A, de Mello Prado R, Habermann E, Lata Tenesaca LF, et al. Silicon mitigates the negative impacts of salt stress in soybean plants. J Sci Food Agric. 2023;103:4360–70. https://doi.org/10.1002/jsfa.12503
  161. 161. Wang Q, Shan C, Zhang P, Zhao W, Zhu G, Sun Y, et al. The combination of nanotechnology and potassium: Applications in agriculture. Environ Sci Pollut Res Int. 2024;31:1–18. https://doi.org/10.1007/s11356-023-31207-y
  162. 162. Hosseinpour A, Haliloglu K, Cinisli KT, Ozkan G, Ozturk HI, Pour-Aboughadareh A, et al. Application of zinc oxide nanoparticles and plant growth promoting bacteria reduces genetic impairment under salt stress in tomato (Solanum lycopersicum L. ‘Linda’). Agriculture. 2020;10(11):521. https://doi.org/10.3390/agriculture10110521
  163. 163. Jalil SU, Ansari MI. Nanoparticles and abiotic stress tolerance in plants: Synthesis, action and signaling mechanisms. In: Khan MIR, Khan NA, editors. Plant signaling molecules: Role and regulation under stressful environments. Cambridge (MA): Academic Press; 2019. p. 549–61. https://doi.org/10.1016/B978-0-12-816451-8.00034-4
  164. 164. Mishra M, Afzal S, Yadav R, Singh NK, Zarbakhsh S. Salinity stress amelioration through selenium and zinc oxide nanoparticles in rice. Sci Rep. 2025;15(1):27554. https://doi.org/10.1038/s41598-025-12106-3
  165. 165. Guo X, Shi Y, Zhu G, Zhou G. Melatonin mitigated salinity stress on alfalfa by improving antioxidant defense and osmoregulation. Agronomy. 2023;13(7):1727. https://doi.org/10.3390/agronomy13071727

Downloads

Download data is not yet available.