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Review Articles

Vol. 13 No. sp2 (2026): Recent Advances in Agriculture

Intercropping for sustainable agriculture: Ecological, agronomical and socioeconomic perspectives

DOI
https://doi.org/10.14719/pst.12408
Submitted
24 October 2025
Published
02-06-2026

Abstract

Intercropping, the simultaneous cultivation of two or more crops on the same land, is a vital strategy for achieving sustainable, resilient and resource-efficient agriculture. By exploiting spatial and temporal complementarities, intercropping enhances biodiversity, nutrient cycling and ecosystem services while minimising environmental impacts. The practice significantly improves land productivity, often reflected by land equivalent ratios (LER) greater than 1.2, indicating superior yield per unit area compared to monocultures frequently reported. Integrating legumes with cereals reduces synthetic nitrogen fertiliser requirements by up to 60 %, thereby lowering greenhouse gas emissions and enhancing soil fertility. Ecologically, intercropping fosters beneficial microbial communities, enhances arbuscular mycorrhizal activity and improves soil organic carbon and nutrient-use efficiency. Agronomically, it suppresses weeds, pests and diseases through diversified canopy structure and allelopathic interactions, while enhancing water-use efficiency and drought resilience in some crop combinations. Socioeconomically, intercropping diversifies farm income, mitigates risks and strengthens smallholder livelihoods. Case studies across Asia, Africa and Europe demonstrate substantial productivity gains, improved soil health and higher carbon sequestration in cereal-legume and agroforestry-based systems. Despite these benefits, challenges such as management complexity, limited mechanisation compatibility and inadequate policy support constrain large-scale adoption. Future directions emphasise breeding for intercropping-suitable cultivars, integrating precision agriculture and digital tools and implementing supportive policy frameworks. As a nature-based solution, intercropping represents a key component cornerstone for sustainable intensification, contributing to climate change mitigation, food security and ecological resilience.

References

  1. 1. Vandermeer J. The ecology of intercropping. Cambridge: Cambridge University Press; 1992.
  2. 2. Brooker RW, Bennett AE, Cong WF, Daniell TJ, George TS, Hallett PD, et al. Improving intercropping: a synthesis of research in agronomy, plant physiology and ecology. New Phytol. 2015;206(1):107–17. https://doi.org/10.1111/nph.13132
  3. 3. Li C, Stomph TJ, Makowski D, Li H, Zhang C, Zhang F, et al. The productive performance of intercropping. Proc Natl Acad Sci U S A. 2023;120:e2201886120. https://doi.org/10.1073/pnas.2201886120
  4. 4. Lithourgidis AS, Dordas CA, Damalas CA, Vlachostergios DN. Annual intercrops: an alternative pathway for sustainable agriculture. Aust J Crop Sci. 2011;5(4):396–410.
  5. 5. Francis CA. Biological efficiency in multiple cropping systems. Adv Agron. 1989;42:1–42. https://doi.org/10.1016/S0065-2113(08)60511-8
  6. 6. Raseduzzaman M, Jensen ES. Does intercropping enhance yield stability in arable crop production? A meta-analysis. Eur J Agron. 2017;91:25–33. https://doi.org/10.1016/j.eja.2017.09.009
  7. 7. Kugbe XJ, Yaro RN, Soyel JK, Kofi ES, Ghaney P. Role of intercropping in modern agriculture and sustainability: a review. Br J Sci. 2018;16(2):67–75. https://www.researchgate.net/publication/335095483_Role_of_Intercropping_in_Modern_Agriculture_and_Sustainability_A_Review
  8. 8. Gliessman SR. Agroecology: the ecology of sustainable food systems. Boca Raton: CRC Press; 2015.
  9. 9. Cassman KG. Ecological intensification of cereal production systems: yield potential, soil quality and precision agriculture. Proc Natl Acad Sci U S A. 1999;96(11):5952–59. https://doi.org/10.1073/pnas.96.11.5952
  10. 10. Pretty J, Benton TG, Bharucha ZP, Dicks LV, Flora CB, Godfray HCJ, et al. Global assessment of agricultural system redesign for sustainable intensification. Nat Sustain. 2018;1(8):441–46. https://doi.org/10.1038/s41893-018-0114-0
  11. 11. Wezel A, Casagrande M, Celette F, Vian JF, Ferrer A, Peigné J. Agroecological practices for sustainable agriculture: a review. Agron Sustain Dev. 2020;40:26. https://doi.org/10.1007/s13593-020-00631-2
  12. 12. Martin-Guay MO, Paquette A, Dupras J, Rivest D. The new green revolution: sustainable intensification of agriculture by intercropping. Sci Total Environ. 2018;615:767–72. https://doi.org/10.1016/j.scitotenv.2017.10.024
  13. 13. Yu Y, Stomph TJ, Makowski D, Zhang L, van der Werf W. A meta-analysis of relative crop yields in cereal/legume mixtures suggests options for optimisation. Field Crops Res. 2021;272:108266. https://doi.org/10.1016/j.fcr.2021.108266
  14. 14. Sileshi G, Mafongoya PL, Kwesiga F, Nkunika P. Termite damage to maize grown in agroforestry systems, traditional fallows and monoculture on nitrogen-limited soils in eastern Zambia. Agric Ecosyst Environ. 2008;125(1-4):189–99.
  15. 15. Agegnehu G, Ghizaw A, Sinebo W. Yield potential and land-use efficiency of barley and faba bean mixed cropping in Ethiopian highlands. Eur J Agron. 2007;28(2):108–17. https://doi.org/10.1016/j.eja.2007.06.002
  16. 16. Bedoussac L, Journet EP, Hauggaard-Nielsen H, Naudin C, Corre-Hellou G, Jensen ES, et al. Ecological principles underlying the increase of productivity achieved by cereal–legume intercrops. Agron Sustain Dev. 2015;35(3):911–35. https://doi.org/10.1007/s13593-014-0277-7
  17. 17. Malézieux E, Crozat Y, Dupraz C, Laurans M, Makowski D, Ozier-Lafontaine H, et al. Mixing plant species in cropping systems: concepts, tools and models. Agron Sustain Dev. 2009;29(1):43–62. https://doi.org/10.1051/agro:2007057
  18. 18. Misra S, Ghosh A. Agriculture paradigm shift: a journey from traditional to modern agriculture. In: Singh K, Ribeiro MC, Calicioglu O, editors. Biodiversity and bioeconomy. Amsterdam: Elsevier; 2024. p. 113–41. https://doi.org/10.1016/B978-0-323-95482-2.00006-7
  19. 19. Food and Agriculture Organisation of the United Nations. Agricultural diversification and intercropping systems. Rome: FAO; 2019.
  20. 20. Mead R, Willey RW. The concept of a land equivalent ratio and the advantages in yields from intercropping. Exp Agric. 1980;16(3):217–28. https://doi.org/10.1017/S0014479700010978
  21. 21. Andersen MK, Nielsen HH, Weiner J, Jensen ES. Competitive dynamics in two and three component intercrops. J Appl Ecol. 2007;44(3):545–51. https://doi.org/10.1111/j.1365-2664.2007.01289.x
  22. 22. Rao MR, Willey RW. Effects of row arrangement in intercropping cowpea with sorghum. Field Crops Res. 1983;7:287–98.
  23. 23. Garrity DP. Agroforestry and the achievement of the Millennium Development Goals. Agrofor Syst. 2004;61:5–17.
  24. 24. Yang Y, Stomph TJ, Makowski D, van der Werf W. Temporal niche differentiation increases the land equivalent ratio of annual intercrops: a meta-analysis. Field Crops Res. 2015;184:133–44. https://doi.org/10.1016/j.fcr.2015.09.010
  25. 25. Cardinale BJ, Duffy JE, Gonzalez A, Hooper DU, Perrings C, Venail P, et al. Biodiversity loss and its impact on humanity. Nature. 2012;486(7401):59–67. https://doi.org/10.1038/nature11148
  26. 26. Jose S. Agroforestry for ecosystem services and environmental benefits: an overview. Agrofor Syst. 2009;76:1–10.
  27. 27. Gaba S, Lescourret F, Boudsocq S, Enjalbert J, Hinsinger P, Journet EP, et al. Multiple cropping systems as drivers for providing multiple ecosystem services: from concepts to design. Agron Sustain Dev. 2015;35:607–23. https://doi.org/10.1007/s13593-014-0272-z
  28. 28. Letourneau DK, Armbrecht I, Rivera BS, Lerma JM, Carmona EJ, Daza MC, et al. Does plant diversity benefit agroecosystems? A synthetic review. Ecol Appl. 2011;21(1):9–21. https://doi.org/10.1890/09-2026.1
  29. 29. Morris RA, Garrity DP. Resource capture and utilisation in intercropping systems. Field Crops Res. 1993;34:319–34. https://doi.org/10.1016/0378-4290(93)90120-C
  30. 30. Richards RA, Watt M, Rebetzke GJ. Root traits and water-use strategies for improved crop performance under drought. J Exp Bot. 2021;72(12):4277–95. https://doi.org/10.1093/jxb/erab155
  31. 31. Zhang H, Li S, Li F, Li L. Intercropping for enhanced water use efficiency and yield stability under drought conditions. Agric Ecosyst Environ. 2022;330:107926. https://doi.org/10.1016/j.agee.2022.107926
  32. 32. Ncube B, Chaplot V, Chimonyo V. Canopy cover and soil microclimate modifications in intercropping systems. Agric Water Manag. 2021;248:106762. https://doi.org/10.1016/j.agwat.2021.106762
  33. 33. Montagnini F, Nair PKR. Carbon sequestration in agroforestry systems. Agrofor Syst. 2004;61(1-3):281–95. https://doi.org/10.1023/B:AGFO.0000029005.92691.79
  34. 34. Dixon RK, Smith J, Guill S. Agroforestry systems: sources and sinks of greenhouse gases? Agrofor Syst. 1995;31(2):99–116. https://doi.org/10.1007/bf00711719
  35. 35. Nair R, Kunhamu TK, Kumar BM. Carbon storage and sequestration in agroforestry systems. Sci Total Environ. 2019;651:1341–57. https://doi.org/10.1016/j.scitotenv.2018.09.312
  36. 36. Cong WF, Hoffland E, Li L, Six J, Sun JH, Bao XG, et al. Intercropping enhances soil carbon and nitrogen. Glob Change Biol. 2015;21(4):1715–26. https://doi.org/10.1111/gcb.12738
  37. 37. Dimande P, Arrobas M, Rodrigues MA. Intercropped maize and cowpea increased the land equivalent ratio. Sustainability. 2024;16(4):1440. https://doi.org/10.3390/su16041440
  38. 38. Yin W, Zhang X, Li Z. Wheat–maize intercropping with reduced tillage and straw retention. Front Plant Sci. 2018;9:1328. https://doi.org/10.3389/fpls.2018.01328
  39. 39. Peichl M, Thevathasan NV, Gordon AM, Huss J. Carbon sequestration potentials in temperate tree-based intercropping systems. Agrofor Syst. 2006;66(3):243–57. https://doi.org/10.1007/s10457-005-0361-8
  40. 40. Yang S, Zhang Y, Li X. Yield performance of maize–soybean intercropping. Field Crops Res. 2024;315:94–106. https://doi.org/10.1016/j.fcr.2024.05.020
  41. 41. Narimani Y, Ghorbani R, Ghanbari A. Cereal-based intercropping systems improve yield advantage. Legume Sci. 2024;6(1):e70002. https://doi.org/10.1002/leg3.70002
  42. 42. Kermah M, Franke AC, Adjei-Nsiah S, Ahiabor BD, Abaidoo RC, Giller KE. Maize-grain legume intercropping for enhanced resource use efficiency. Field Crops Res. 2017;211:75–87. https://doi.org/10.1016/j.fcr.2017.07.008
  43. 43. Zhou L, Su L, Zhao H, Zhao T, Zheng Y, Tang L. Maize/soybean intercropping improves yield stability and sustainability. Agronomy. 2024;14:1222. https://doi.org/10.3390/agronomy14061222
  44. 44. Nasar J, Ahmad M, Gitari H, Tang L, Chen Y, Zhou XB. Maize/soybean intercropping increases nutrient uptake and modifies soil characteristics. BMC Plant Biol. 2024;24:350. https://doi.org/10.1186/s12870-024-05061-0
  45. 45. Gidey T, Berhe DH, Birhane E. Intercropping maize with faba bean improves yield and soil fertility. Scientifica. 2024;2024:2552695. https://doi.org/10.1155/2024/2552695
  46. 46. Pankou C, Lithourgidis A, Menexes G, Dordas C. Importance of cultivar selection in wheat-pea intercropping systems. Agronomy. 2022;12(10):2367. https://doi.org/10.3390/agronomy12102367
  47. 47. Pambuka GT, Kinge TR, Ghosh S, Cason ED, Nyaga MM, Gryzenhout M. Pea-wheat intercrops in low-input conditions. Eur J Agron. 2012;40:1–8. https://doi.org/10.1016/j.eja.2012.01.010
  48. 48. Šenk M, Simić M, Milojković-Opsenica D. Common millet and soybean intercropping with bio-fertiliser. Front Nutr. 2023;10:1267928. https://doi.org/10.3389/fnut.2023.1267928
  49. 49. Liu X, Zhao. Maize–soybean intercropping improved maize growth traits by increasing soil nutrients and reducing plant pathogen abundance. Front Microbiol. 2023. https://doi.org/10.3389/fmicb.2023.1290825
  50. 50. Pambuka GT, Kinge TR, Ghosh S, Cason ED, Nyaga MM, Gryzenhout M. Plant and soil core mycobiomes in a two-year sorghum-legume intercropping system. Microorganisms. 2022;10(10):2079. https://doi.org/10.3390/microorganisms10102079
  51. 51. Government of India. National Mission for Sustainable Agriculture (NMSA) under the National Action Plan on Climate Change. https://nmsa.dac.gov.in/
  52. 52. European Commission. Agriculture and Rural Development: Common Agricultural Policy overview. https://agriculture.ec.europa.eu/common-agricultural-policy/cap-overview_en
  53. 53. Agegnehu G, Ghizaw A, Sinebo W. Crop productivity and land-use efficiency of a teff/faba bean mixed cropping system. Exp Agric. 2006;42:495–504. https://doi.org/10.1017/S0014479706003863
  54. 54. Feder G, Murgai R, Quizon JB. The acquisition and diffusion of knowledge in farmer field schools, Indonesia. J Agric Econ. 2011;62(1):13–38. https://doi.org/10.1111/j.1477-9552.2010.00280.x

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